Yuto makes his appearance on the DenLiner, and the Owner mentions that the ZeroLiner is another train that can travel through time and Sakurai may be its Kamen Rider, but he believes the ZeroLiner had been destroyed. After a losing fight with the Jelly Imagin, Yuto arrives and transforms into Kamen Rider Zeronos. He fights the Jelly Imagin as Altair Form, and uses Deneb to slow the Jelly Imagin down until Deneb can no longer hold on. Yuto chastises his Imagin, and together they become Vega Form and destroy the Jelly Imagin.